The shortcomings of present population genetic theory are discussed as they (pertain) to problems of speciation, extinction and the evolution of genetic systems. Which of the following can replace the word in brackets?

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The shortcomings of present population genetic theory are discussed as they (pertain) to problems of speciation, extinction and the evolution of genetic systems. Which of the following can replace the word in brackets?

A.presume
B.attribute
C.relate
D.trace
正确答案C
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